Whenever a creature you control leaves the battlefield, put a +1/+1 counter on Traxian Hoplite.

Noncreature spells you cast cost {1} less to cast.

At the beginning of combat on your turn, discard a card, then draw a card. If you control three or more Shrines, instead discard a card, scry 2, then draw a card.

At the beginning of combat on your turn, put a +1/+1 counter on target creature you control. If you control three or more Shrines, instead put two +1/+1 counters on that creature instead.

Deathtouch

Karda's Emmisary is a 5/5 Illusion enchantment creature as long as you control three or more enchantments.

At the beginning of combat on your turn, create a 0/0 white Army creature token, then put a +1/+1 counter on it. If you control three or more Shrines, instead create that token, then put a +1/+1 counter on each Army you control.

Ward {2}
Imposing Serpent enters the battlefield with X +1/+1 counters on it.

At the beginning of combat on your turn, up to one target creature you control has base power and toughness 3/3 until end of turn. If you control three or more Shrines, that creature has base power and toughness 6/6 until end of turn instead.

Flying
When Anvilwrought Crane dies, create a Weapon token.

When Chromatic Hart enters the battlefield, you may exile target card from a graveyard.
{T}: Add one mana of any color.

This spell costs {X} less to cast, where X is your greatest devotion to a single color.
Menace

Whenever Implacable Shrine-Guard attacks, for each color, do the following if your devotion to that color is three or greater.
{W} — Create a 1/1 white Human creature token.
{U} — Draw a card.
{B} — Each opponent loses 2 life.
{R} — Create a Treasure token.
{G} — You gain 3 life.

{1}, {T}: Add one mana of any color for each Shrine you control.

When The Smoking Strand enters the battlefield, you may search your library for a planeswalker card, reveal it, put it into your hand, then shuffle.
Planeswalkers you control have “+1: Draw a card.”

When Soul-Guide Lantern enters the battlefield, exile target card from a graveyard.
{T}, Sacrifice Soul-Guide Lantern: Exile each opponent's graveyard.
{1}, {T}: Sacrifice Soul-Guide Lantern: Draw a card.

{T}: Add one mana of any color. Put a tide counter on Tidelock Pearl.
{1}{U}, {T}, Sacrifice Tidelock Pearl: Search your library for a Kraken, Leviathan, Octopus, or Serpent card with mana value less than or equal to the number of tide counters on Tidelock Pearl, put it onto the battlefield, then shuffle.

Barren Ashlands enters the battlefield tapped.
{T}: Add {B} or {R}.
{5}{B/R}: Barren Ashlands becomes a 3/3 black and red Lizard creature with first strike until end of turn. It's still a land. This ability costs {X} less to activate, where X is your devotion to black and red. Activate only once each turn.

Buried Colossus enters the battlefield tapped.
{T}: Add {G} or {W}.
{8}{G/W}: Buried Colossus becomes a 8/8 green and white Cyclops creature until end of turn. It's still a land. This ability costs {X} less to activate, where X is your devotion to green and white. Activate only once each turn.

Ephemeral Mire enters the battlefield tapped.
{T}: Add {U} or {B}.
{4}{U/B}: Until end of turn, Ephemeral Mire becomes a 3/2 black and blue Horror creature until end of turn and can't be blocked this turn. It's still a land. This ability costs {X} less to activate, where X is your devotion to black and blue. Activate only once each turn.

{T}: Add {C}.
{T}: Add one mana of any color. Spend this mana only to cast Shrine spells or activate abilities of Shrines.

Gates to the Underworld enters the battlefield tapped.
{T}: Add {W} or {B}.
{5}{W/B}: Gates to the Underworld becomes a 3/4 white and black Construct creature with lifelink until end of turn. It's still a land. This ability costs {X} less to activate, where X is your devotion to white and black. Activate only once each turn.

Hundred-Handed Pillars enters the battlefield tapped.
{T}: Add {W} or {U}.
{7}{W/U}: Hundred-Handed Pillars becomes a 4/6 white and blue Giant creature with vigilance until end of turn. It's still a land. This ability costs {X} less to activate, where X is your devotion to white and blue. Activate only once each turn.

Miasmic Cove enters the battlefield tapped.
{T}: Add {B} or {G}.
{4}{B/G}: Miasmic Cove becomes a 2/3 black and green Snake creature with deathtouch until end of turn. It's still a land. This ability costs {X} less to activate, where X is your devotion to black and green. Activate only once each turn.

Nessian Woods enters the battlefield tapped.
{T}: Add {R} or {G}.
{6}{R/G}: Nessian Woods becomes a 5/4 red and green Dryad creature with trample until end of turn. It's still a land. This ability costs {X} less to activate, where X is your devotion to red and green. Activate only once each turn.